Almost certainly issue with the sockets for the bulbs, exacerbated by swapping them around (and you likely had the initial right rear one blown) - it is very easy to mash the contacts in the socket shorting them out (or pushing them so far in they no longer make contact with the contact); this is particularly common when using the after-market orange bulbs and clear lenses

So - combination of bulbs and socket contacts - use your known working bulb/location to validate each position and each bulb.

(to check each bulb you can also take a piece of wire stripped at both ends - hold one end against the body of the bulb between your finger and thumb - touch the contact on the base of the bulb to one of the battery terminals being VERY careful to only touch the center contact of the bulb and not the main body - touch the other end of the stripped wire to the opposite terminal)
Do you have a picture of how the bulb socket should look without a bulb in it?
 
Not sure if I can piggy back this thread or should start my own. I bought mine over the winter its a 2008 not the touring package, but has had a lot of mods done to it including a full led upgrade to all the lighting. Went on a 4 hour trip and about half way through they just stopped, horn and brake lights all still work but no blinkers on any of them. I was looking into it and most things said relay which led me here but now I'm not sure I even have one.
 
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