I'm confused by what you mean with the first statement. 2.45kOhms @22 deg??? How do you know that resistance value for that temp is good? Do you have some kind of legend you can reference?
The biggest issue I see is no rad fan function. You have to find out why. Obviously it functions since you can manually activate it. I'm not sure exactly how the fan is activated but I imagine the temp sensor is monitored by the ECM which appears to ground the fan relay, turning it on, when the 101c temp is reached. Since you already swapped fan relays, check that temp sensor closely (I'm guessing it is the problem). I'm assuming that the wiring and connectors are good and plugged in properly at the sensor, relay and ECM because you can manually turn the fan on with Tune ECU.